I only realized that after finding them together in an archive. Took me a while to figure out how the story goes too. The anon takes an Alice family and abuses them to death, then he takes a next family, and then another, and now this koyukkuri. Right?
Now look at the very last panel, this one above all comments. There is a saying that when someone dies, in last moments he sees all his life scrolled in his inner sight. Effect surrounding picture is quite fitting for this. And so, I presume, we see past when once-seemed-so-kind-anon picked up future mother Alice when she was yet a lone koyukkuri (lost her family?) saving her from certain death by it. And of course she was deeply touched and loved Anon for this.
the main idea of the story is :the anon's beloved pet yukkuri reimu were raped to death by a gang of yukkuri alices, which turned all his devoted love to reimu into a twisted hatred of alice. He breeds them carefully and then torture them by cruel means.